Paul Nolan

Address: Department of Biology, The Citadel
Phone: 843.953.7076


Ph.D., 2002, Auburn University
M.S., 1993, The University of California at Davis

Research Interests

  • Behavioral ecology
  • Disease ecology
  • Ornithology
  • Avian habitat selection

Current and planned research projects

  • signal content and function of ornamental colors in penguins
  • demography of avian susceptibility to West Nile Virus on a coastal plain
  • human impacts on colonially nesting seabirds
  • avian use of stormwater ponds


  • Nolan, Paul M., F. Stephen Dobson, Marion Nicolaus, Tim J. Karels, Kevin J. McGraw, and Pierre Jouventin.  2010.  Mutual mate choice for colorful traits in king penguins. Ethology 116:635-644.
  • Dobson, F. Stephen, Paul M. Nolan, Marion Nicolaus, Catherine Bajzak, Anne-Sophie Coquel, and Pierre Jouventin. 2008. Comparison of color and body condition between early- and late-breeding King Penguins. Ethology 114:925-933.
  • Viera, Vanessa M., Paul M. Nolan, Steeve D. Côté, Pierre Jouventin, René Groscolas.  2008.  Is territory defence related to plumage ornaments in the king penguin Aptenodytes patagonicus?  Ethology 114:146-153.
  • Nicolaus, Marion, Céline Le Bohec,  Paul M. Nolan, Michel Gauthier-Clerc, Yvon Le Maho, Jan Komdeur, and Pierre Jouventin.  2007.  Coloured and UV ornaments reveal age in king penguins.  Polar Biology 31:53-61.
  • Nolan, Paul M., Birgitta Dresp, F. Stephen Dobson, and Pierre Jouventin.  2006.  Immunocompetence is signalled by ornamental color in king penguins,Aptenodytes patagonicus.  Evolutionary Ecology Research 8:1325-1332.
  • Nolan, Paul M., Sharon R. Roberts, and Geoffrey E. Hill.  2004.  Effects ofMycoplasma gallisepticum on reproductive success in house finches.  Avian Diseases 48:879-885.
  • Nolan, Paul M., Geoffrey E. Hill and Andrew M. Stoehr.  1998.  Sex, size, and plumage redness predict house finch survival in an epidemic.  Proceedings, Royal Society of London, Series B 265:961-965.