Andrew Clark

College of Charleston

Address: RITA 123, Department of Biology, Charleston, SC 29424
Personal Website:


Ph.D., 2009, University of California, Irvine

Research Interests

  • Comparative biomechanics of animal feeding behaviors
  • Mechanical properties of biological and synthetic materials
  • Animal movement and locomotion

Current and planned research projects

  • Material properties of the integument from elongate fishes
  • Morphology, tensile mechanics, and puncture resistance in cockroach cuticle
  • Functional morphology of jawless biting and knotting behaviors in hagfish


  • Haney, William A., Clark, Andrew J. and Theodore A. Uyeno. 2020. Characterization of body knotting behavior used for escape in a diversity of hagfishes. Journal of Zoology. 310: 261-272.
  • Clubb, Benjamin L., Clark, Andrew J. and Theodore A. Uyeno. 2019. Powering the hagfish “bite”: the functional morphology of the retractor complex of two hagfish feeding apparatuses. Journal of Morphology280: 827-840.
  • Lees, Lauren E., Krueger-Hadfield, Stacy A., Clark, Andrew J., Duermit, Elizabeth A., Sotka, Erik E., Murren, Courtney J. 2018. Nonnative Gracilaria vermiculophylla tetrasporophytes are more difficult to debranch and are less nutritious than gametophytes. Journal of Phycology. 54: 471-482.
  • Schumacher, Erika L., Owens, Brad D., Uyeno, Theodore A., Clark, Andrew J., and Joshua S. Reece. 2017. No support for Heincke’s Law in hagfish (Myxinidae): lack of an association between body size and the depth of species occurrence. Journal of Fish Biology. 91: 545-557.
  • Clark, Andrew J., Crawford, Callie H., King, Brooke D., Demas, Andrew M. and Theodore A. Uyeno. 2016. Material properties of hagfish skin with insights into knotting behaviors. Biological Bulletin. 230: 243–256.
  • Uyeno, Theodore A. and Andrew J. Clark. 2015. Muscle articulations: flexible jaw joints made of soft tissues. Integrative and Comparative Biology. 55(2): 193-204.
  • Clark, Andrew J. and Jeffrey D. Triblehorn. 2014. Mechanical properties of the cuticles of three cockroach species that differ in their wind-evoked escape behavior. PeerJ. 2:e501
  • Clark, Andrew J. and Adam P. Summers. 2012. Ontogenetic scaling of the morphology and biomechanics of the feeding apparatus in the Pacific hagfish, Eptatretus stoutii. Journal of Fish Biology. 80: 86-99.
  • Clark, Andrew J. and Timothy E. Higham. 2011. Slipping, sliding, and stability: locomotor strategies for overcoming low-friction surfaces. Journal of Experimental Biology. 214: 1369- 1378.
  • Fuller, Patrick O., Higham, Timothy E., and Andrew J. Clark. 2011. Posture, speed, and habitat structure: Three-dimensional hindlimb kinematics of two species of padless geckos. Zoology. 114: 104-112.
  • Clark, Andrew J., Maravilla, Erick J., and Adam Summers. 2010. A soft origin for a forceful bite: motor patterns of the feeding musculature in Atlantic hagfish, Myxine glutinosaZoology. 113: 259-268. (COVER)